Canada geese head tossing

The Canada geese with a broken wing is now in the company of his partner in the park, who didn't leave with the flock in early March to their breeding grounds. He managed to retain two other individuals for a while - maybe offspring from previous years, as Canada Geese have long term family ties - but they ended up leaving. This morning, somebody started feeding the birds on an end of the pond, and I saw the pair of Canada swimming towards the feeding point. Broken Wing was leading his partner. He kept tossing his head every few seconds, and after a few tries I managed to capture the behaviour (above). It felt like a 'hurry up, follow me!' beckoning to female. Indeed, Canada Geese often communicate using head movements. I have covered the pre-flight head tossing, which is accompanied by much calling. When on land, and during the flightless phase of chick rearing during the moulting period, head-tossing by itself indicates readiness to walk or swim and is used especially when they are very motivated to move. Head tossing is very conspicuous due to the white cheek patch, and this is often directed to family members. Individuals head-toss when stationary, appearing to signal to other family members to follow them, including females to young goslings, and the signal is continued during movement. The direction at which the head is tossed is also informative, indicating the direction of movement. Even young goslings - who don't have the white patch - start head tossing in the first day after hatching, signalling their intention to move to other foraging patch. In a situation of threat, e.g. when a predator is located, this visual signalling allows parents to quietly direct their goslings to cover, avoiding attracting unwanted attention by acoustic signals.

More information

Jeffrey M. Black and James H. Barrow, Jr. (1985) Visual signalling in Canada geese for the coordination of family units. Wildfowl, 36, 35-41. Available here.

Canada Geese taking off decisions

This morning, the 60 strong flock of Canada Geese that seemed to have roosted in the park were restless. There were continuous loud grunts and honks and, after a crescendo in which more and more individuals joined in the calling, part of the flock took off in a coordinated way. The vocalising geese were making a decision, with individuals deciding to join or not a party leading the departure, maybe to quieter feeding grounds. It is unclear if the individuals felt hungry or if the several dog walkers had made then nervous (or both!).
  Flocking birds have to travel together to stay in a group to search for new feeding or roosting patches, to avoid predation or to migrate. When the individuals differ in their motivations to move away (some might be hungrier than others, for example), there might be a period in which an a consensus is reached about leaving or staying involving communication of the individuals intentions. An individual that is not hungry might change its mind and join in if most of the flock appear willing to move, so as not to become isolated.
  Dennis Raveling, in a paper in 1969, reported on the behaviour before flock taking off in Canada Geese. He observed a large flock in their natural range, in which 77 geese had been marked and radio tracked, including 10 families. Flock departure was preceded by a ceremony, with the neck stretched, there are quick head tossing movements with the bill pointing up and repeatedly, and the white head patch conspicuously displayed - communicating an intention to fly. Geese often spread and flap their wings and start to walk in the intended direction of flight for a few steps (this video illustrate this behavior). Ganders (adult male geese) were more successful at recruiting his family than any other family members, as a shorted time elapsed from his initiation of head-tossing until the family took flight, although all family members initiated head tossing at some point. In a couple of occasions when an excited immature took flight but the rest of the family did not follow, it flew in a circle and returned with the family shortly.
 Other than the cohesive function of the head-toss ceremony for family members, group vocalisations and wing flapping serves to synchronise the whole flock. The coordination of the flock during take off included the presence of an invisible 'starting line': individuals run until they arrived at the same position at which the individual before them had just taken off (instead of taking off where they were). Raveling hypothesized that the contrast between the white tail coverts and the black tail served as a signal to optimise the position of individuals during flight, quite important in such large birds. Families tended to keep together during flight and their vocalisations then changed from grunting to a more trumpet-like honking.

More information

Dennis G. Raveling (1969). Preflight and Flight Behavior of Canada Geese The Auk, 86, 671-681 DOI: 10.2307/4083454